Summary

Did you know?

This is the only member of this genus and family that occurs in North America. The genus is confined to the New World (Philbrick and Novelo 2004). The genus name Podostemum comes from the greek words for foot and stamen and is in reference to the stamens appearing "raised on a stalk by the side of the ovary" (Fernald 1970). The specific epithet ceratophyllum means horn-leaf (Fernald 1970) and may be in reference to the narrow stiff leaves of this species.

State Ranking Justification

There are 16 known extant populations and 8 populations which have not been seen in recent years and are considered historical. Many of the populations are clustered along particular rivers. Habitat is somewhat limited for this species but certainly not rare.

Short-term Trends

Fifteen out of the 16 known extant populations have first been documented in the past 30 years. This would seem to point to an increase in this species but since this species is often overlooked, resembles algae, and grows in habitat that make it difficult to detect, it is equally possible that these "new" populations were simply overlooked in the past. Overall, short term trends are not clear but probably are at least stable.

Long-term Trends

No population of this species is known to have become extirpated. Eight populations have not been seen in over 25 years. These populations may still be extant, as targeted surveys at these populations have not been conducted. The habitat for this species may have decreased some in the past 100+ years as rivers and streams were developed and/or polluted. Overall long term trends are not clear.

Conservation and Management

Threats

Some of the streams where this species occurs may be used for a water source. This development could result in a change to the stream that would make the habitat less suitable for riverweed. Other populations are threatened by water quality deterioration as a result of development, agriculture, and run off from a sewage treatment plant (New York Natural Heritage Program 2007). Negative impacts on rivers including siltation, nutrient pollution, and dam building have been considered threats to this species (Quiroz et al. 1997 in Philbrick and Novelo 2004).

Conservation Strategies and Management Practices

Water quality of the rivers where this species occurs, should be protected.

Research Needs

Surveys to historical populations are needed to determine if these populations are still extant.

Habitat

Habitat

Riverweed grows submerged (up to 75+ cm deep) to seasonally exposed on cobbles and bedrock substrate in fast flowing, relatively large streams or rivers. It often occurs in rapids (New York Natural Heritage Program 2008). Fast-moving streams, especially rapids (Crow and Hellquist 2000). In rapidly flowing water (Gleason and Cronquist 1991). On rocks in streams (Fernald 1970).

Associated Ecological Communities

• Confined river (guide)
  The aquatic community of relatively large, fast flowing sections of streams with a moderate to gentle gradient.
• Rocky headwater stream (guide)
  The aquatic community of a small- to moderate-sized perennial rocky stream typically with a moderate to steep gradient, and cold water that flows over eroded bedrock, boulders, or cobbles in the area where a stream system originates.

Associated Species

• Cladophora sp.
• Platyhypnidium riparioides

Range

New York State Distribution

In New York, this species occurs in northern New York, including the Adirondacks and southeastern New York, including the Catskills with one population reported from Long Island and one from central New York (New York Natural Heritage Program 2008).

Global Distribution

This species occurs from Nova Scotia and New Brunswick, west to Quebec and southeastern Ontario, and south to Georgia, Louisiana, Arkansas, and Oklahoma. It is disjunct in the Dominican Republic, and Honduras (Philbrick and Novelo 2004).

Identification Comments

General Description

Riverweed is an olive-green, aquatic plant of firm texture that grows in streams and rivers attached to rocks. It resembles seaweed, alga, or moss and attaches to rocks by fleshy disks. The stems are 2-30 cm long and often branched. The leaves are very narrow and divided into numerous linear lobes. The flowers are small, not showy, and scattered along the stems (Fernald 1970, Gleason and Cronquist 1991, Philbrick and Novelo 2004).

Best Life Stage for Proper Identification

It is easiest to identify this plant when it is in flower or fruit although vegetative material, when not confused with alga or moss, is also relatively easy to identify.

Similar Species

This species is perhaps superficially similar to alga and moss but can be distinguished from both by its true although minute flowers, its firm texture, fleshy disks (which attach to the substrate), and stipule-like appendages at the bases of the leaves. The latter three characters will also help separate this species from other aquatic vascular plants.

Best Time to See

Flowers and/or fruits are present from mid-June through early October. Since this species often grows submerged in rapids and fast moving water it can be difficult to spot. Therefore, the best time to survey for this species is between mid-June and early October and when water levels are low.

Riverweed Images

Taxonomy

Riverweed
Podostemum ceratophyllum Michx.

• Kingdom Plantae
  • Phylum Anthophyta
    • Class Dicotyledoneae (Dicots)
      • Order Podostemales
        • Family Podostemaceae

Additional Common Names

• Horn-leaved riverweed
• River-moss
• Threadfoot

Synonyms

• Podostemon ceratophyllum Michaux [orthographic variant]

Comments on the Classification

When Michaux (1803) described this genus he used the spelling "Podostemum" in the text and then in a plate illustrating Podostemum ceratophyllum he used the spelling "Podostemon". It appears that Michaux's Podostemon was in error (Sprague 1933) and the family Podostemaceae (based on Podostemum) has been conserved as the correct spelling (McNeill et al. 2006). Podostemon is therefore considered an orthographic variant and is to be replaced by the correct spelling Podostemum.

Additional Resources

Best Identification Reference

Gleason, Henry A. and A. Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York. 910 pp.

Other References

Crow, Garrett E. and C. Barre Hellquist. 2000. Aquatic and Wetland Plants of Northeastern North America: A revised and enlarged edition of Norman C. Fassett's a Manual of Aquatic Plants. Volume One: Pteridophytes, Gymnosperms, and Angiosperms: Dicotyledons. The University of Wisconsin Press. Madison, Wisconsin. 536 Pages.

Fernald, M.L. 1950. Gray's manual of botany. 8th edition. D. Van Nostrand, New York. 1632 pp.

Gleason, Henry A. 1952. The New Britton and Brown Illustrated Flora of the Northeastern United States and Canada.

Haines, Arthur and Thomas F. Vining. 1998. Flora of Maine. A Manual for Identification of Native and Naturalized Vascular Plants of Maine.

Holmgren, Noel. 1998. The Illustrated Companion to Gleason and Cronquist's Manual. Illustrations of the Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York.

McNeill, J., F.R. Barrie, H.M. Burdet, V. Demoulin, D.J. Hawksworth, K. Marhold, D.H. Nicolson, J. Prado, P.C. Silva, J.E. Skog, J.H. Wiersema, and N.J. Turland (Editors). 2006. International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventh International Botanical Congress Vienna, Austria, July 2005. A.R.G. Gantner Velag, Ruggell, Liechtenstein.

Michaux, A. 1803. Flora boreali-americana, sistens caracteres plantarum quas in America septentrionali collegit et detexit Andreas Michaux. Apud Fratres Levrault, Paris and Strasbourg, France.

New York Natural Heritage Program. 2010. Biotics database. New York Natural Heritage Program. New York State Department of Environmental Conservation. Albany, NY.

New York Natural Heritage Program. 2023. New York Natural Heritage Program Databases. Albany, NY.

Philbrick, C.T. and A. Novelo R. 2004 Monograph of Podostemum (Podostemaceae). Systematic Botany Monographs 70: 1-106.

Reschke, Carol. 1990. Ecological communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation. Latham, NY. 96 pp. plus xi.

Rhoads, Ann F. and Timothy A. Block. 2000. The Plants of Pennsylvania, an Illustrated Manual. University of Pennsylvania Press, Philadelphia, PA.

Sprague, T.A. 1933. Podostemaceae or Podostemonaceae. Bulletin of Miscellaneous Information (Royal Gardens, Kew) 46: 46.

Weakley, A. S. 2007. Flora of the Carolinas, Virginia, Georgia, and surrounding areas. Working draft of 11 January 2007. University of North Carolina Herbarium (NCU), North Carolina Botanical Garden, University of North Carolina at Chapel Hill. Online. Available: http://www.herbarium.unc.edu/flora.htm (accessed 2007).

Weldy, T. and D. Werier. 2010. New York flora atlas. [S.M. Landry, K.N. Campbell, and L.D. Mabe (original application development), Florida Center for Community Design and Research http://www.fccdr.usf.edu/. University of South Florida http://www.usf.edu/]. New York Flora Association http://newyork.plantatlas.usf.edu/, Albany, New York

Weldy, Troy W. and David Werier. 2005. New York Flora Atlas. [S.M. Landry, K.N. Campbell, and L.D. Mabe (original application development), Florida Center for Community Design and Research. University of South Florida]. New York Flora Association, Albany, NY. Available on the web at (http://newyork.plantatlas.usf.edu/).

Links

• Google Images
• NatureServer Explorer
• New York Flora Atlas
• USDA Plants Database

About This Guide

Information for this guide was last updated on: January 22, 2008

Please cite this page as:
New York Natural Heritage Program. 2023. Online Conservation Guide for Podostemum ceratophyllum. Available from: https://guides.nynhp.org/riverweed/. Accessed September 24, 2023.