Jeffersonia diphylla in flower Stephen M. Young

Jeffersonia diphylla in flower
Stephen M. Young

Class
Dicotyledoneae (Dicots)
Family
Berberidaceae (Barberry Family)
State Protection
Threatened
Listed as Threatened by New York State: likely to become Endangered in the foreseeable future. For animals, taking, importation, transportation, or possession is prohibited, except under license or permit. For plants, removal or damage without the consent of the landowner is prohibited.
Federal Protection
Not Listed
State Conservation Status Rank
S2
Imperiled in New York - Very vulnerable to disappearing from New York due to rarity or other factors; typically 6 to 20 populations or locations in New York, very few individuals, very restricted range, few remaining acres (or miles of stream), and/or steep declines.
Global Conservation Status Rank
G5
Secure globally - Common in the world; widespread and abundant (but may be rare in some parts of its range).

Summary

Did you know?

Twinleaf is one of only two species in the genus Jeffersonia. The other species, Jeffersonia dubia, grows in Mongolia. Benjamin Barton, a Philadelphia botanist of the 18th-century, was so impressed by Thomas Jefferson's knowledge of natural history he named this plants after him. Twinleaf is said to have been used for treating rheumatism but it can be poisonous if not prepared correctly.

State Ranking Justification

There are 15 populations in the state with two thirds of them containing hundreds to thousands of plants. There are 23 historical populations, many of which have not been searched, and more populations are expected to be found. While the short term trends are stable, logging and invasion by exotic species may negatively affect long term trends.

Short-term Trends

Based on limited survey work, most populations have remained stable in the last two decades.

Long-term Trends

As a plant that thrives in ravine areas, this plant has received limited protection as ravines are difficult to log and develop. Based on this habitat type, population numbers are presumably similar to early settlement periods. Since New York is near the edge of its range, this is one of the plants that has likely always been rare within New York. The invasion of garlic mustard does present some concerns for its long-term stability. More research is needed here.

Conservation and Management

Threats

A significant immediate threat is the invasion of its habitat by garlic mustard. Continued logging at and near some of its populations has allowed garlic mustard to invade the area. It is unclear however if twinleaf is able to withstand these invasions because of its clonal nature. At a minimum, garlic mustard is likely to limit the recruitment of new plants from seed.

Conservation Strategies and Management Practices

A large buffer (minimum suggested buffer is 100 meters although this could increase due to other factors such as slope and nearby activities) should be established around each population to protect the woodland slope environment from disturbance such as logging or ATV use. These disturbances often allow for the establishment of garlic mustard and other invasives which may outcompete twinleaf.

Research Needs

Research detailing the competitive interactions between garlic mustard and twinleaf would allow us to better determine the threat posed by garlic mustard and other invasive species.

Habitat

Habitat

This plant is typically encountered near the bases of slopes that were created by a stream cutting down through the bedrock. These stream cuts expose the calcareous bedrock and soils preferred by this plant. Most often the plants are found near the tow of the slope and could be present within the floodplain zone of these small streams. A few populations are known from more upland settings associated with calcareous soils, but these populations tend to be much smaller (New York Natural Heritage Program 2006). Moist woods, usually on calcareous soils (Rhoads and Black 2000). Rich moist woods to semi-open rocky slopes and outcrops, usually over limestone or other calcareous rocks, at elevations of 100-800 meters (FNA 1997). Rich woods, preferring calcareous soil (Gleason and Cronquist 1991). Rich deciduous woods, including floodplains and well-drained slopes (Voss 1985). Woods (Fernald 1970).

Associated Ecological Communities

  • Calcareous talus slope woodland* (guide)
    An open or closed canopy community that occurs on talus slopes composed of calcareous bedrock such as limestone or dolomite. The soils are usually moist and loamy; there may be numerous rock outcrops. * probable association but not confirmed.
  • Floodplain forest (guide)
    A hardwood forest that occurs on mineral soils on low terraces of river floodplains and river deltas. These sites are characterized by their flood regime; low areas are annually flooded in spring, and high areas are flooded irregularly.
  • Limestone woodland (guide)
    A woodland that occurs on shallow soils over limestone bedrock in non-alvar settings, and usually includes numerous rock outcrops. There are usually several codominant trees, although one species may become dominant in any one stand.
  • Maple-basswood rich mesic forest (guide)
    A species rich hardwood forest that typically occurs on well-drained, moist soils of circumneutral pH. Rich herbs are predominant in the ground layer and are usually correlated with calcareous bedrock, although bedrock does not have to be exposed. The dominant trees are sugar maple, basswood, and white ash.
  • Oak-tulip tree forest* (guide)
    A hardwood forest that occurs on moist, well-drained sites in southeastern New York. The dominant trees include a mixture of five or more of the following: red oak, tulip tree, American beech, black birch, red maple, scarlet oak, black oak, and white oak. * probable association but not confirmed.
  • Rich mesophytic forest (guide)
    A hardwood or mixed forest that resembles the mixed mesophytic forests of the Allegheny Plateau south of New York but is less diverse. It occurs on rich, fine-textured, well-drained soils that are favorable for the dominance of a wide variety of tree species. A canopy with a relatively large number of codominant trees characterizes this forest. Canopy codominants include five or more of the following species: red oak, red maple, white ash, American beech, sugar maple, black cherry, cucumber tree, and black birch.

Associated Species

  • Acer saccharum (sugar maple)
  • Actaea pachypoda (white baneberry, doll's-eyes)
  • Alliaria petiolata (garlic mustard)
  • Allium tricoccum var. tricoccum (common wild leek)
  • Arisaema triphyllum ssp. triphyllum (common jack-in-the-pulpit)
  • Asarum canadense (wild ginger)
  • Cardamine diphylla (two-leaved toothwort)
  • Carex careyana (Carey's sedge)
  • Carex jamesii (James's sedge)
  • Carex tetanica (rigid sedge)
  • Carpinus caroliniana ssp. virginiana (musclewood, ironwood, American hornbeam)
  • Carya cordiformis (bitternut hickory)
  • Caulophyllum thalictroides (blue cohosh, late blue cohosh)
  • Circaea lutetiana ssp. canadensis
  • Dryopteris intermedia (evergreen wood fern, fancy wood fern, common wood fern)
  • Equisetum hyemale var. affine
  • Erythronium albidum (white trout-lily)
  • Erythronium americanum
  • Fagus grandifolia (American beech)
  • Floerkea proserpinacoides (false mermaid-weed)
  • Fraxinus americana (white ash)
  • Geranium maculatum (wild geranium)
  • Juglans nigra (black walnut)
  • Lindera benzoin (spicebush)
  • Lonicera tatarica (Tartarian honeysuckle)
  • Ostrya virginiana (hop hornbeam, ironwood)
  • Parthenocissus quinquefolia (Virginia-creeper)
  • Podophyllum peltatum (may-apple)
  • Prunus serotina
  • Prunus virginiana
  • Quercus rubra (northern red oak)
  • Rhus aromatica
  • Sanguinaria canadensis (bloodroot)
  • Solidago caesia
  • Solidago flexicaulis (zig-zag goldenrod)
  • Symplocarpus foetidus (skunk-cabbage)
  • Thalictrum thalictroides (rue-anemone)
  • Tilia americana var. americana (American basswood)
  • Toxicodendron radicans ssp. radicans (eastern poison-ivy)
  • Trillium grandiflorum (white trillium)
  • Viola affinis (LeConte's violet)

Range

New York State Distribution

This spring ephemeral is mainly limited to calcareous woods of central and western New York. We do not know if the few Hudson Valley populations represent native populations, planted populations, or escapes from cultivation. There is a possibility this plant will appear elsewhere in the state as a planted population or escape from cultivation.

Global Distribution

This plant of rich, usually calcareous, woods ranges from western New York, southern Ontario, Michigan, Wisconsin, and southeastern Minnesota, southward to the mountain areas of North Carolina, Georgia, and Alabama.

Best Places to See

  • Sonyea State Forest (Livingston County)
  • Powder Mills Park (Monroe County)
  • Oakwood Cemetery (Rensselaer County)
  • Sampson State Park (Seneca County)

Identification Comments

General Description

Twinleaf is an early spring wildflower that flowers before the forest trees have leafed-out. When it flowers, the stems are about 8 inches high with small two-lobed leaves that look like butterfly wings. Later in the season the expanded leaves grow taller, up to 20 inches, and obscure the stems. The showy white flowers are solitary at the top of each stem, have eight separate petals and are about 1/2" to 1 inch wide with yellow stamens in the middle. An individual plant may hold its flower for a few days. The fruit is walnut-shaped, smooth and shiny green, and often sticks out above the leaves. When the fruit is mature it splits around the top and opens like a lid, releasing the seeds.

Identifying Characteristics

This is a perennial spring wildflower. The white flowers have eight distinct petals, although four drop off very quickly. The leaves are on a long petiole up to 50 cm tall. These leaves are deeply cleft into two large symmetrical lobes. The fruit capsule is 2-3 cm long and shiny green to yellow in color. This spring plant typically grows in clustered colonies where populations often include 100s to 1000s of individuals.

Best Life Stage for Proper Identification

This is a very unique looking plant that can be identified in leaf, flower, or fruit; although it is easiest to locate while in flower. A photograph should suffice as evidence/verification.

Similar Species

This is a very easy to identify spring wildflower that is not easily confused with other wildflowers. Some may initially confuse the flower with bloodroot (Sanguinaria canadensis), but the leaves of these plants are very different.

Best Time to See

This spring wildflower flowers from mid-April to mid-May. Fruits begin to appear in May with seeds dispersing by mid-summer. Vegetative leaves persist until the first frosts, although the leaves are in poor shape by late summer. Since this plant is easiest to locate while flowering, surveys are most effective when conducted mid-April to mid-May; however, this plant can be identified vegetatively so surveys could continue until late summer.

  • Vegetative
  • Flowering
  • Fruiting

The time of year you would expect to find Twinleaf vegetative, flowering, and fruiting in New York.

Twinleaf Images

Taxonomy

Twinleaf
Jeffersonia diphylla (L.) Pers.

  • Kingdom Plantae
    • Phylum Anthophyta
      • Class Dicotyledoneae (Dicots)
        • Order Ranunculales
          • Family Berberidaceae (Barberry Family)

Additional Common Names

  • Rheumatism Root

Additional Resources

Best Identification Reference

Gleason, Henry A. and A. Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York. 910 pp.

Other References

Fernald, M.L. 1950. Gray's manual of botany. 8th edition. D. Van Nostrand, New York. 1632 pp.

Holmgren, Noel. 1998. The Illustrated Companion to Gleason and Cronquist's Manual. Illustrations of the Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York.

House, Homer D. 1924. Annotated list of the ferns and flowering plants of New York State. New York State Museum Bulletin 254:1-758.

Mitchell, Richard S. and Gordon C. Tucker. 1997. Revised Checklist of New York State Plants. Contributions to a Flora of New York State. Checklist IV. Bulletin No. 490. New York State Museum. Albany, NY. 400 pp.

New York Natural Heritage Program. 2010. Biotics database. New York Natural Heritage Program. New York State Department of Environmental Conservation. Albany, NY.

New York Natural Heritage Program. 2019. New York Natural Heritage Program Databases. Albany, NY.

Rhoads, Ann F. and Timothy A. Block. 2000. The Plants of Pennsylvania, an Illustrated Manual. University of Pennsylvania Press, Philadelphia, PA.

Voss, E.G. 1985. Michigan flora. Part II. Dicotyledons. Cranbrook Institute of Science and University of Michigan Herbarium. Ann Arbor, Michigan. 1212 pp.

Weldy, T. and D. Werier. 2010. New York flora atlas. [S.M. Landry, K.N. Campbell, and L.D. Mabe (original application development), Florida Center for Community Design and Research http://www.fccdr.usf.edu/. University of South Florida http://www.usf.edu/]. New York Flora Association http://newyork.plantatlas.usf.edu/, Albany, New York

Links

About This Guide

Information for this guide was last updated on: January 14, 2009

Please cite this page as:
New York Natural Heritage Program. 2019. Online Conservation Guide for Jeffersonia diphylla. Available from: https://guides.nynhp.org/twinleaf/. Accessed May 26, 2019.

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